Robert Weladji

Assistant Professor,
Concordia University

Department of Biology
Institution page
Personal page


Postdoctoral research in Life History Strategies and Evolutionary Ecology (2003-2005) at: The Norwegian University of Life Sciences, The Centre for Ecological and Evolutionary Synthesis at the University of Oslo, and Laboratoire de Biométrie et Biologie Évolutive at Université Claude Bernard Lyon 1 in France.
PhD in Ecology, Norwegian University of Life Sciences (2003)
MSc in Management of Natural Resources, Agricultural University of Norway
BSc in Wildlife and Forestry, University of Dschang, Cameroon


My research interest is rather broad and includes natural resource management, population ecology, behavioral and evolutionary ecology, wildlife conservation and management, terrestrial ecosystems ecology and grazing ecology. During the last years, I have worked mainly with (1) the influence of environmental conditions (including global climatic change) on life history and population dynamics of ungulates, mainly large herbivores (e.g. reindeer/caribou, sheep); (2) life history strategies and evolutionary ecology of mammals, mainly reindeer/caribou and sheep, and (3) behavioral ecology (rutting behavior, mating strategies, social interactions, etc.). Together with colleagues from Norway and Finland, I am increasingly being interested in understanding sexual strategies (mating strategies, mate choice, costs of sexual reproduction, etc.) and the influence of the male composition (e.g. age structure, sex ratio) on population ecology of cervids. The major challenge of my research is to integrate all this knowledge so as to promote sustainable conservation and management of the biodiversity (fauna and flora). Indeed, I am also interested in (4) management and conservation of biodiversity in the tropics and plan to establish a long term research program in Africa. To achieve all these, I can count on collaborators in Canada, Norway, Finland, France, Uganda, South Africa and Cameroon, but also on the contribution from the students.

QCBS students

Isaac Blaise DJOKO

(Start: 2017, End: 2021)
Interactions People-Protected Areas: The case of Campo-Ma’an National Park, Southern Cameroon

Louis Lazure

(Start: 2017, End: 2022)
Behavioural impact of human presence on mammal mesopredators in protected areas
Personal page

Amélie Paoli

(Start: 2015)
Breeding phenology and calving synchrony of reindeer in relation to climatic variability
Personal page

Laura Puch

(Start: 2020)
The role of male vocalisations in sexual selection in reindeer (Rangifer tarandus)
Personal page

Julienne Therese Bonoan

(Start: 2018)
Social Network Analysis of female reindeer Rangifer tarandus: Do females associate due to relatedness?
Personal page

Catherine Pelletier

Other (co-)supervisor: Patrick Paré, Zoo de Granby
(Start: 2017)
Impact of ultrasounds and visitors on captive animal behaviour and welfare

Previous students (since 2009)

Guillaume Body

(Start: 2010, End: 2013)
Reindeer Rangifer tarandus female mating tactics: dealing with the fission-fusion dynamic system
Personal page

Sacha Engelhardt

(Start: 2012, End: 2016)
Suckling and allosuckling in reindeer (Rangifer tarandus): the evolution of sociality and cooperative breeding.

Franco Alo

(Start: 2014, End: 2016)
The influence of operational sex ratio (OSR) on the intensity of competition and male mating tactics in reindeer (Rangifer tarandus)
Personal page

Julie Beaudin-Judd

(Start: 2015, End: 2017)
Impact of open exhibit concept on captive animal behaviour and welfare, and on visitor perception
Personal page

Jeffrey Driscoll

(Start: 2016, End: 2018)
The relationship between operational sex ratio (OSR) and the intensity of competition in reindeer (Rangifer tarandus L.): the effect of male age

Caitlin Finlay

(Start: 2015, End: 2017)
The importance of social structure and environment on breeding success in captive flamingoes (Phoenicopterus ruber)
Personal page

Issac Hebert

(Start: 2010, End: 2012)
Characterizing core caribou areas and their change in space with forestry development in Newfoundland

Leon L'Italien

(Start: 2006, End: 2010)
Females' mate choice in reindeer: effects of age and social rank of males

Karine Laflamme-Mayer

(Start: 2008, End: 2010)
Influence of Environmental Conditions on Sex Allocation in the Black Rhinoceros (Diceros bicornis minor) Population of Mkhuze Game Reserve, South Africa (2008-2010)

Sara McCarthy

(Start: 2008, End: 2010)
Landscape Composition and Configuration Influences Woodland Caribou Calf Recruitment in Newfoundland

Natalka Melnycky

(Start: 2010, End: 2012)
Allometric growth in secondary sex traits: tradeoffs when allocating resources and a reflection of individual quality in reindeer (Rangifer tarandus)

C-Jae Morden

(Start: 2008, End: 2010)
Faecal sampling as a non-invasive population monitoring and management method for reindeer and caribou

Erica Tennenhouse

(Start: 2008, End: 2010)
Reproductive effort in rutting dominant male reindeer: the influence of mating group composition, age, and rutting period

Examples of species studied

Rangifer tarandus
Elephas maximus
Ovis aries
Bos grunniens


87- Seasonal reproduction of northernmost endangered forest musk deer (Moschus berezovskii) in China and the synchronization with climatic conditions
Wang J , Zheng Q , Xia C , Li Y , Zhou M , Sheng Y , Weladji R B , Meng X ,
2021 Czech Journal of Animal Science (Date Added: 2021-03-29)

86- Zoo soundscape: Daily variation of low-to-high-frequency sounds
Pelletier C , Weladji R B , Lazure L , Paré P ,
2020 Zoo Biology (Date Added: 2020-08-09)

85- Biodiversity and carbon sequestration potential in two types of tropical rainforest, Cameroon
Kabelong Banoho L , Zapfack L , Weladji R B , Chimi Djomo C , Nyako M C , Nasang J M , Madountsap Tagnang N , Tabue Mbobda R B ,
2020 Acta Oecologica (Date Added: 2020-04-30)

84- Early-life conditions determine the between-individual heterogeneity in plasticity of calving date in reindeer
Paoli A , Weladji R B , Holand ,
2019 Journal of Animal Ecology (Date Added: 2019-11-02)

83- Temporal pattern of parturition in captive Alpine musk deer (Moschus sifanicus)
Zhao Y , Wang J , Li Y , Zhou M , Weladji R B , Bonoan J T , Meng X ,
2019 Biologia (Date Added: 2019-07-04)

82- Summer habitat selection of reindeer (rangifer tarandus) governs on the unprotected forest and human interface in China
Wang J , Wang P , Aryal A , Meng X , Weladji R B ,
2018 Ekologia Bratislava (Date Added: 2018-12-03)

81- Floristic diversity and carbon stocks in the periphery of Deng–Deng National Park, Eastern Cameroon
Kabelong Banoho L P R , Zapfack L , Weladji R B , Chimi Djomo C , Nyako M C , Bocko Y E , Essono D M , Nasang J M , Madountsap Tagnang N , Memvi Abessolo C I , Mvondo Sakouma K R , Mamadou Souahibou F , Palla F J S , Peguy T K , Jiagho R , Kenmou T L , Choumele Kana Jumo U A , Anyam Yi Andjik B A , Tabue Mbobda R B ,
2018 Journal of Forestry Research (Date Added: 2018-11-27)

80- Winter and spring climatic conditions influence timing and synchrony of calving in reindeer
Paoli A , Weladji R B , Holand ,
2018 PLoS ONE (Date Added: 2018-05-04)

78- Temporal variation in the operational sex ratio and male mating behaviours in reindeer (Rangifer tarandus)
Weladji R B , Body G , Holand ,
2017 Behavioural Processes (Date Added: 2017-05-09)

76- Allonursing in reindeer, Rangifer tarandus: a test of the kin-selection hypothesis
Engelhardt, SC, Weladji, RB, Holand, , Røed, KH, Nieminen, M
2016 Journal of Mammalogy (Date Added: 2016-09-02)

75- Endangered wild yak (Bos grunniens) in the Tibetan plateau and adjacent regions: Population size, distribution, conservation perspectives and its relation to the domestic subspecies
Shi, Q, Guo, Y, Engelhardt, SC, Weladji, RB, Zhou, Y, Long, M, Meng, X
2016 Journal for Nature Conservation (Date Added: 2016-05-09)

74- Allosuckling in reindeer (Rangifer tarandus): A test of the improved nutrition and compensation hypotheses
Engelhardt, SC, Weladji, RB, Holand, , Nieminen, M
2016 Mammalian Biology - Zeitschrift für Säugetierkunde (Date Added: 2016-04-13)

73- Les gorilles du Parc National de Lobéké (Cameroun): interactions avec les populations locales et implications pour la conservation
Tsakem, S, Tchamba, M, Weladji, R
2015 International Journal of Biological and Chemical Sciences (Date Added: 2016-07-06)

72-Abondance et Distribution des Grands Singes et des Activites Humaines dans le Parc National de Campo Ma’an (Sud Cameroun)
M N Tchamba, PCFH
2015 Tropicultura ( Date Added: 2016-06-15)

71- Foraging competition in larger groups overrides harassment avoidance benefits in female reindeer (Rangifer tarandus)
Uccheddu, S, Body, G, Weladji, RB, Holand, O, Nieminen, M
2015 Oecologia (Date Added: 2015-11-11)

70- Measuring variation in the frequency of group fission and fusion from continuous monitoring of group sizes
Body, G, Weladji, RB, Holand, , Nieminen, M
2015 Journal of Mammalogy (Date Added: 2015-10-23)

69- Evidence of Reciprocal Allonursing in Reindeer, Rangifer tarandus
Engelhardt, SC, Weladji, RB, Holand, , Røed, KH, Nieminen, M
2015 Ethology (Date Added: 2015-02-04)

68-Les Gorilles du Parc National de Lobéké, (Cameroun) : Utilisation de l’habitat et Implications pour la conservation.
S C Tsakem, MNT&RBW
2014 Parcs et reserves ( Date Added: 2016-07-06)

67- Highly competitive reindeer males control female behavior during the rut
Body, G, Weladji, RB, Holand, O, Nieminen, M
2014 PloS one (Date Added: 2016-07-06)

66-Plant biomass density as an indicator of food supply for elephants (Loxodonta africana) in Waza National Park, Cameroon
M N Tchamba, RBW
2014 Tropical Conservation Science ( Date Added: 2016-06-15)

65- Allosuckling in reindeer (Rangifer tarandus): Milk-theft, mismothering or kin selection?
Engelhardt, SC, Weladji, RB, Holand, , de Rioja, CM, Ehmann, RK, Nieminen, M
2014 Behavioural Processes (Date Added: 2014-10-16)

64- Effects of males’ presence on female behaviour during the rut
Djakovi?, N, Holand, , Hovland, AL, Weladji, RB, Røed, KH, Nieminen, M
2014 Ethology Ecology & Evolution (Date Added: 2014-07-17)

63- Fission-fusion group dynamics in reindeer reveal an increase of cohesiveness at the beginning of the peak rut
Body, G, Weladji, RB, Holand, , Nieminen, M
2014 acta ethologica (Date Added: 2014-07-08)

62- The Effect of Zoo Visitors on Activity Patterns of Captive African Herbivores
T Kalioujny, RBW
2013 Journal of Biodiversity Management & Forestry (Date Added: 2016-06-15)

61- Scaling of antler size in reindeer ( Rangifer tarandus ): sexual dimorphism and variability in resource allocation
Melnycky, NA, Weladji, RB, Holand, , Nieminen, M
2013 Journal of Mammalogy (Date Added: 2014-01-06)

60- The use of coniferous forests and cutovers by Newfoundland woodland caribou
Hébert, I, Weladji, RB
2013 Forest Ecology and Management (Date Added: 2013-01-11)

58- The recursive model as a new approach to validate and monitor activity sensors
Body, G, Weladji, RB, Holand,
2012 Behavioral Ecology and Sociobiology (Date Added: 2012-10-09)

57- Timing of Reproductive Effort Differs between Young and Old Dominant Male Reindeer
Tennenhouse, EM, Weladji, RB, Holand, , Nieminen, M
2012 Annales Zoologici Fennici (Date Added: 2012-09-18)

56- Induced orphaning reveals post-weaning maternal care in reindeer
Holand, , Weladji, RB, Mysterud, A, Røed, KH, Reimers, E, Nieminen, M
2012 European Journal of Wildlife Research (Date Added: 2012-09-18)

55- Movement and occurrence of two elephant herds in a human-dominated landscape, the Bénoué Wildlife Conservation Area, Cameroon
Granados, A, Weladji, RB, Loomis, MR
2012 Tropical Conservation Science (Date Added: 2012-08-03)

54- Mating Group Size and Stability in Reindeer Rangifer tarandus: The Effects of Male Characteristics, Sex Ratio and Male Age Structure
L'Italien, L, Weladji, RB, Holand, , Røed, KH, Nieminen, M, Côté, SD
2012 Ethology (Date Added: 2012-07-16)

53- Human–Elephant Conflict Around Bénoué National Park, Cameroon: Influence on Local Attitudes and Implications for Conservation
Granados, A, Weladji, RB
2012 Human Dimensions of Wildlife (Date Added: 2012-04-20)

52- Association patterns and kinship in female reindeer (Rangifer tarandus) during rut
Djakovi?, N, Holand, , Hovland, AL, Røed, KH, Weladji, RB, Fjeldstad, E, Nieminen, M
2011 acta ethologica (Date Added: 2012-01-09)

51- Influence of environmental conditions on sex allocation in the black rhinoceros population of Mkhuze Game Reserve, South Africa
Weladji, RB, Laflamme-Mayer, K
2011 African Journal of Ecology (Date Added: 2011-12-16)

50- Woodland caribou calf recruitment in relation to calving/post-calving landscape composition
McCarthy, SC, Weladji, RB, Doucet, C, Saunders, P
2011 Rangifer (Date Added: 2011-08-30)

49- Fecal hormones as a non-invasive population monitoring method for reindeer
Morden, C-JC, Weladji, RB, Ropstad, E, Dahl, E, Holand, , Mastromonaco, G, Nieminen, M
2011 The Journal of Wildlife Management (Date Added: 2011-08-26)

48- Mating group composition influences somatic costs and activity in rutting dominant male reindeer (Rangifer tarandus)
Tennenhouse, EM, Weladji, RB, Holand, O, Røed, KH, Nieminen, M
2011 Behavioral Ecology and Sociobiology (Date Added: 2011-04-14)

47- Scale-dependent effects of summer density on autumn mass in reindeer
Holand, , Anders, A, Weladji, RB
2010 Rangifer (Date Added: 2011-08-30)

46- Comparative response of Rangifer tarandus and other northern ungulates to climatic variability
Weladji, RB, Klein, DR, Holand, , Mysterud, A
2010 Rangifer (Date Added: 2011-08-26)

45- Role of traditional enclosures on the diversity of herbaceous vegetation in a semi-arid rangeland, southern Ethiopia
Angassa, A, Oba, G, Treydte, AC, Weladji, RB
2010 Livestock Research for Rural Development (Date Added: 2011-04-14)

44- Age-specific changes in different components of reproductive output in female reindeer: Terminal allocation or senescence?
Weladji, RB, Holand, O, Gaillard, J-M, Yoccoz, NG, Mysterud, A, Nieminen, M, Stenseth, NC
2010 Oecologia (Date Added: 2011-04-14)

43-Embracing Ethnoveterinary Knowledge Diffusion in Karamoja, Uganda: A Strategy to Strengthen
Knaute, D, Weladji, R
2009 Sustainability in Karamoja ? : rethinking the terms of global sustainability in a crisis region of Africa ( Date Added: 2011-08-25)

42- Healer-driven ethnoveterinary knowledge diffusion among semi-nomadic pastoralists in Karamoja, Uganda
Gradé, J, Weladji, R, Tabuti, J, Van Damme, P
2009 Afrika Focus (Date Added: 2011-08-25)

41- Variation in calf body mass in migratory caribou: The role of habitat, climate, and movements
Couturier, S, Côté, SD, Otto, RD, Weladji, RB, Huot, J
2009 Journal of Mammalogy (Date Added: 2011-04-14)

40- Scaling effects of proximate desertification drivers on soil nutrients in northeastern Tanzania
Oba, G, Weladji, RB, Msangameno, DJ, Kaitira, LM, Stave, J
2008 Journal of Arid Environments (Date Added: 2011-04-14)

39- Heterogeneity in individual quality overrides costs of reproduction in female reindeer
Weladji, RB, Loison, A, Gaillard, J-M, Holand, , Mysterud, A, Yoccoz, NG, Nieminen, M, Stenseth, NC
2008 Oecologia (Date Added: 2011-04-14)

38- Feedback effects of chronic browsing on life-history traits of a large herbivore
Anouk Simard, M, Côté, SD, Weladji, RB, Huot, J
2008 Journal of Animal Ecology (Date Added: 2011-04-14)

37- Un siècle de broutement intensif sur l’île d’Anticosti
Simard, A, Côté, SD, Weladji, RB, de Bellefeuille, S, Huot, J
2007 Naturaliste Canadien (Date Added: 2012-03-13)

36- No evidence of inbreeding avoidance in a polygynous ungulate: The reindeer (Rangifer tarandus)
Holand, , Askim, KR, Røed, KH, Weladji, RB, Gjøstein, H, Nieminen, M
2007 Biology Letters (Date Added: 2011-04-14)

34- Good reindeer mothers live longer and become better in raising offspring.
Weladji, RB, Gaillard, JM, Yoccoz, NG, Holand, O, Mysterud, A, Loison, A, Nieminen, M, Stenseth, NC
2006 Proceedings of the Royal Society B: Biological Sciences (Date Added: 2011-04-14)

33- The role of area enclosures and fallow age in the restoration of plant diversity in northern Ethiopia
Abebe, MH, Oba, G, Angassa, A, Weladji, RB
2006 African Journal of Ecology (Date Added: 2011-04-14)

32- Male age structure influences females' mass change during rut in a polygynous ungulate: The reindeer (Rangifer tarandus)
Holand, , Weladji, RB, Røed, KH, Gjøstein, H, Kumpula, J, Gaillard, J-M, Smith, ME, Nieminen, M
2006 Behavioral Ecology and Sociobiology (Date Added: 2011-04-14)

31- Constraints to projecting the effects of climate change on mammals
Berteaux, D, Humphries, MM, Krebs, CJ, Lima, M, McAdam, AG, Pettorelli, N, Réale, D, Saitoh, T, Tkadlec, E, Weladji, RB, Stenseth, NC
2006 Climate Research (Date Added: 2011-04-14)

30- Adaptive adjustment of offspring sex ratio and maternal reproductive effort in an iteroparous mammal
Holand, , Mysterud, A, Røed, KH, Coulson, T, Gjøstein, H, Weladji, RB, Nieminen, M
2006 Proceedings of the Royal Society B: Biological Sciences (Date Added: 2011-04-14)

29- Sexual dimorphism and intercorhort variation in reindeer calf antler length is associated with density and weather
Weladji, RB, Holand, , Steinheim, G, Colman, JE, Gjøstein, H, Kosmo, A
2005 Oecologia (Date Added: 2011-04-14)

28- Effects of anthropogenic fire history on savanna vegetation in northeastern Namibia
Sheuyange, A, Oba, G, Weladji, RB
2005 Journal of Environmental Management (Date Added: 2011-04-14)

27- Dry season diets and habitat use of sympatric Asian elephants (Elephas maximus) and greater one-horned rhinoceros (Rhinocerus unicornis) in Nepal
Steinheim, G, Wegge, P, Fjellstad, JI, Jnawali, SR, Weladji, RB
2005 Journal of Zoology (Date Added: 2011-04-14)

26- Differences in choice of diet between sheep breeds grazing mountain pastures in Norway
Steinheim, G, Nordheim, LA, Weladji, RB, Gordon, IJ, Ådnøy, T, Holand,
2005 Acta Agriculturae Scandinavica - Section A: Animal Science (Date Added: 2011-04-14)

25- The relative role of winter and spring conditions: Linking climate and landscape-scale plant phenology to alpine reindeer body mass
Pettorelli, N, Weladji, RB, Holand, , Mysterud, A, Breie, H, Stenseth, NC
2005 Biology Letters (Date Added: 2011-04-14)

24- Effect of calf stimulation on milk ejection in reindeer (Rangifer tarandus)
Gjøstein, H, Holand, , Bolstad, T, Weladji, RB
2004 Rangifer (Date Added: 2011-08-30)

23- Social rank in female reindeer (Rangifer tarandus): Effects of body mass, antler size and age
Holand, , Gjøstein, H, Losvar, A, Kumpula, J, Smith, ME, Røed, KH, Nieminen, M, Weladji, RB
2004 Journal of Zoology (Date Added: 2011-04-14)

22- Chimpanzee diet and habitat selection in the Budongo Forest Reserve, Uganda
Tweheyo, M, Lye, KA, Weladji, RB
2004 Forest Ecology and Management (Date Added: 2011-04-14)

21- Reproductive effort in relation to maternal social rank in reindeer (Rangifer tarandus)
Holand, , Weladji, RB, Gjøstein, H, Kumpula, J, Smith, ME, Nieminen, M, Røed, KH
2004 Behavioral Ecology and Sociobiology (Date Added: 2011-04-14)

20- Milk production and composition in reindeer (Rangifer tarandus): Effect of lactational stage
Gjøstein, H, Holand, O, Weladji, RB
2004 Comparative Biochemistry and Physiology - A Molecular and Integrative Physiology (Date Added: 2011-04-14)

19- Climatic variability and effects on ungulate body weight: The case of domestic sheep
Steinheim, G, Weladji, RB, Skogan, T, Ådnøy, T, Skjelvåg, AO, Holand,
2004 Annales Zoologici Fennici (Date Added: 2011-04-14)

18- Use of indigenous ecological knowledge of the Maasai pastoralists for assessing rangeland biodiversity in Tanzania
Mapinduzi, AL, Oba, G, Weladji, RB, Colman, JE
2003 African Journal of Ecology (Date Added: 2011-04-14)

17- Use of climatic data to assess the effect of insect harassment on the autumn weight of reindeer (Rangifer tarandus) calves
Weladji, RB, Holand, , Almøy, T
2003 Journal of Zoology (Date Added: 2011-04-14)

16- Stakeholder attitudes towards wildlife policy and the Bénoué Wildlife Conservation Area, North Cameroon
Weladji, RB, Moe, SR, Vedeld, P
2003 Environmental Conservation (Date Added: 2011-04-14)

15- Sex ratio variation in reindeer Rangifer tarandus: A test of the extrinsic modification hypothesis
Weladji, RB, Holand, O
2003 Wildlife Biology (Date Added: 2011-04-14)

14- Scale-dependent effects of grazing on rangeland degradation in northern Kenya: A test of equilibrium and non-equilibrium hypothesis
Oba, G, Weladji, RB, Lusigi, WJ, Stenseth, NC
2003 Land Degradation and Development (Date Added: 2011-04-14)

13- Effect of pre-oestrus body weight on lambing success in primiparous sheep (Ovis aries)
Weladji, R, Steinheim, G, Post, E, Holand, , Ådnøy, T
2003 Acta Agriculturae Scandinavica - Section A: Animal Science (Date Added: 2011-04-14)

12- An assessment of restoration of biodiversity in degraded high mountain grazing lands in northern Ethiopia
Asefa, DT, Oba, G, Weladji, RB, Colman, JE
2003 Land Degradation and Development (Date Added: 2011-04-14)

11- Temporal patterns of juvenile body weight variability in sympatric reindeer and sheep
Weladji, RB, Steinheim, G, Holand, , Moe, SR, Almøy, T, Ådnøy, T
2003 Annales Zoologici Fennici (Date Added: 2011-04-14)

10- Sex-specific preweaning maternal care in reindeer (Rangifer tarandus t.)
Weladji, RB, Holand, , Steinheim, G, Lenvik, D
2003 Behavioral Ecology and Sociobiology (Date Added: 2011-04-14)

9- Maternal age and offspring sex ratio variation in reindeer (Rangifer tarandus)
Weladji, RB, Holand, , Yoccoz, NG, Lenvik, D
2003 Annales Zoologici Fennici (Date Added: 2011-04-14)

7- Digestive tract anatomy of Norwegian sheep: Difference between breeds
Stenheim, G, Nordheim, LA, Weladji, RB, Holand, , Ådnøy, T
2003 Acta Agriculturae Scandinavica - Section A: Animal Science (Date Added: 2011-04-14)

5- Effect of "owners" selection strategies on autumn weight in reindeer (Rangifer t. tarandus) calves
Weladji, RB, Holand, , Steinheim, G, Hansen, H
2002 Rangifer (Date Added: 2011-08-30)

3- Age-related reproductive effort in reindeer (Rangifer tarandus): Evidence of senescence
Weladji, RB, Mysterud, A, Holand, , Lenvik, D
2002 Oecologia (Date Added: 2011-04-14)

2- Impacts of shifting agriculture on a floodplain woodland regeneration in dryland, Kenya
Oba, G, Stenseth, NC, Weladji, RB
2002 Agriculture, Ecosystems and Environment (Date Added: 2011-04-14)

1-Gestion participative des aires protégées dans la province du Nord-Cameroun
Koulagna, K, Weladji, R
1996 Edition SNV, Cameroun ( Date Added: 2011-08-26)